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Graduate Courses


Graduate Courses


B5800 Microbial Ecology: Interrelations of microorganisms with other organisms and the abiotic environment. 2 LECT, 4 LAB HR./WK., OR A FIELDTRIP; 4 CR.

V0503: Evolution: Study of the mechanism and processes of evolution. Theory, laboratory experimental results and the phenomena found in natural populations are described and discussed in relation to population genetics, speciation and megaevolution. Prereq: course in genetics, vertebrate or invertebrate zoology, botany or permission of instructor. 3 LECT. HR./WK.; 3 CR.

V0507: Fossil Record: 3 LECT. HR./WK.; 3 CR.

V0603: Principles of Systematics: Lecture and discussion sections involving general principles of biological systematics, including fundamentals of nomenclature, phylogenetic theory, character analysis, and their use of relevant computer algorithms. Readings from the primary literature as well as text sources will be emphasized. 3 LECT. HR./WK., PLUS CONF.; 4 CR.

V0611/12: Systematics and Evolution of Insects and Spiders: Lectures emphasize basic knowledge and recent advances in the systematics, biogeography, morphology, behavior, and paleontology of these arthropods. Major Families of the world will be emphasized. Labs involve fieldwork, personal collections, identifications, techniques, and small research projects. 2 LECT. HR./WK., 2 CR.; 4 LAB HR./WK., 2 CR.

V0733/34: Zoology and Phylogeny of Chordata (Mammals): Lecture and Laboratory on the origin, adaptive radiation, morphology, ecology and systematics of mammals. Discussion of the reptilemammal transition emphasizing the fundamental characters of teeth, ear structure and tarsal bones. Survey of mammalian orders and practical work in laboratory on living families and local species, including field methods and preparation of specimens for study. Lecture and laboratory are integrated; the course cannot be taken in separate parts. Prereq: course in vertebrate comparative anatomy. 2 LECT. HR./WK., 2 CR.; 5 LAB HR./WK., 2 CR.

V0743/44: Zoology and Phylogeny of Chordata (Birds): Lecture and laboratory on special topics in the evolution of birds. Prereq: permission of the instructor. 2 LECT. HR./WK., 2 CR.; 4 LAB HR./WK., 2 CR.

V0901/02: Population Genetics: Lecture and laboratory on the Hardy-Weinberg law, gene pools, gene frequencies, and gene migration. Prereqs: a course in genetics, a course in organic chemistry. 3 LECT. HR./WK., 3 CR.; 6 LAB HR./WK., 3 CR.

 V6003/04: Community Ecology: Lecture and laboratory on Structural attributes, growth, and regulation of plant and animal communities. Prereq: a course in either ecology or field biology. 3 LECT. HR./WK., 3 CR.; 6 LAB HR./WK., 3 CR.

V6005/06: Population Ecology: Lecture, laboratory. An analysis of the structure and dynamics of plant and animal populations. Topics include density, growth, regulation, fluctuation of numbers, niche, dispersal systems, dispersion patterns, demographic techniques, and interactions between populations. 3 LECT. HR./WK., 3 CR.; 6 LAB HR./WK., 3 CR.

V6101/02: Ecology of Marine Plankton: Lecture, laboratory. Biology, productivity and trophic dynamics of marine plankton. Prereqs: a course in biological oceanography, a course in organic chemistry. 3 LECT. HR./WK., 3 CR.; 6 LAB. HR./WK., 3 CR.

V6200/01: Physiological Ecology: Comparative study of physiological mechanisms important in adaptation to different environments. Focus is on the biotic and abiotic factors in the habitat and the adaptations that determine an animal's ability to survive. Prereq: a course in physiology or cell biology. 3 LECT. HR./WK., 3 CR.; 6 LAB HR./WK., 3 CR.

V6701/02: Biology of Fishes: Lecture and Laboratory. 3 LECT. HR./WK., 3 CR.; 6 LAB. HR./WK., 3 CR.

V9001: Seminar in Evolution: Topics relating to the general subject of evolution. 2 HR./WK., PLUS CONF.; 3 CR.

V9006: Seminar in Ecology: The conservation ecology seminar will focus on genetic problems and implications of wildlife management programs. Prereq: permission of the instructor. 3 HR./WK.; 3 CR.

V9012: Seminar in Zoogeography: Special topics are discussed and reviewed. Prereq: permission of the instructor. 2 HR./WK., PLUS CONF.; 3 CR.

V9030: Seminar in Ecology, Evolution, and Behavior AMNH: (Alternate weeks). 2 HR./WK.; 1 CR.



B7400: Cell Microsurgery: Lecture, laboratory. The course is designed to teach the principles and operation of micromanipulators for experimental cell research. Prereqs: undergraduate background in cell biology or cell physiology or equivalent and permission of instructor. 6 LAB. HR./WK.; 3 CR.

V0005: Genetics: Prokaryotic and eukaryotic genetics; organization of DNA, replication, repair, mutagenesis, recombination, control of gene expression, genetic engineering and molecular techniques. Prereq: undergraduate genetics and molecular biology or biochemistry. 4 LECT. HR./WK.; 4 CR.

V0103: Microbial Genetics: Microbial genetic systems will be examined with respect to their contributions to the understanding of molecular mechanisms of recombination, repair of genetic material, and regulation of gene expression. Emphasis will be placed on the procedures and the role of the new biotechnology. Prereq: V0005 or equivalent. 3 LECT. HR./WK.; 3 CR.
V0803: Molecular Evolution: Principles of evolution at the level of DNA  and proteins; gene families, concerted evolution  of genes, codon bias, and other  genetic processes will be discussed. 3  LECT. HR./WK.; 3 CR.   

V1401: Cell Biology: Cells will be studied with special emphasis  placed on organization, molecular structure/  function relationships of organelles,  and energetics and metabolism. Prereqs: a  course in organic chemistry and a course  in biochemistry or permission of the  instructor. 4 LECT. HR./WK.; 4 CR.  



V2101: Animal Physiology I:  This course includes an introduction to  physiology; cell structure and function of  specialized cells; cellular metabolism; thermodynamics,  kinetics and energetics;  nutrition, feeding, digestion and metabolism;  respiration; circulation; temperature;  and locomotion. 4 LECT. HR./WK.; 4 CR.

V2102: Animal Physiology II: Physiological control mechanisms, including regulation of water and ions; rhythms and physiological states; growth, maturation and aging; self-recognition systems and adaptation to environmental extremes. 4 LECT. HR./WK.; 4 CR. 

V2301: Neuroscience I: This course and V2302 comprise an introduction to the neurosciences. The first semester covers neuronal cell biology (structure and trophic functions), cellular neurophysiology (membrane physiology, action potentials), synapses (neurotransmitters and physiology), neuroendocrine mechanisms, local neuronal circuits (integrative anatomical organization of the vertebrate nervous system). The second semester will cover the sensory and motor systems, as well as neurochemical pathways. Receptor physiology and information processing are discussed in brief for the somatosensory system and in detail for the visual and auditory systems. The vertebrate motor systems are discussed with respect to spinal, supraspinal and cortical mechanisms. Prereq: students with no background in physiology must meet instructor before course begins. 4 LECT. HR./WK.; 4 CR.

V2302: Neuroscience II: Building upon materials covered in Neuroscience I, this course provides an introduction to sensory processing for several sensory systems, outlines the important developmental processes with a specific focus on the functional differentiation of the brain. 3 LECT., 1 CONF. HR./WK.; 4 CR.

V2403: Animal Behavior I: Lectures and discussions of selected major areas in modern animal behavior research. Included among the topics are conceptual issues in methodology, orientation and navigation, and development of behavior. A modern eclectic approach is emphasized. Students read and lead discussion of papers from the original literature. Prereq: an undergraduate course in animal behavior/ comparative psychology or permission of the instructor. 3 LECT. HR./WK.; 3 CR.

V2404: Biological Basis of Animal Behavior: Laboratory Apprenticeship training in the laboratories of behavioral scientists. Prereqs: undergraduate laboratory course in animal behavior (Bio 46000 or equivalent) and permission of the instructor. 6 LAB HR./WK.; 3 CR.

V2407: Animal Behavior II: Lecture and discussion of the selected major areas in modern behavioral research. Topics include behavioral genetics, the evolution of behavior, and behavioral ecology. Prereq: an undergraduate course in animal behavior, evolution or ecology. 3 HR./WK; 3 CR.

V2505: Animal Communication V2601/02: Comparative Animal Physiology: Study of the physiological mechanisms common to a wide variety of animals. Focus is on the underlying physiochemical processes and functions of organ systems. Prereq: a course in physiology or cell biology. 3 LECT. HR./WK., 3 CR.; 6 LAB HR./WK., 3 CR.

V9101: Colloquium in Ecology, Evolution and Behavior AMNH: (Alternate weeks). 2 LECT. HR./WK.; 1 CR.


B9700: Special Topics: 3 LECT HR./WK., 3 CR.; 6 LAB HR./WK., 3 CR.

V4103/04: Radiation Biology: Lecture and laboratory on a broad unified coverage of the effects of ionizing radiation and the application of tracer techniques in biological systems at the molecular, cellular, organ, organism, and community levels. Pre- or coreq: Cell Physiology. 2 LECT. HR./WK., 2 CR.; 4 LAB HR./WK., 2 CR.

V7200: Biological Electron Microscopy: Preparation of materials and their examination by means of Transmission and Scanning electron microscopes. Techniques include methods of fixation and embedment for TEM; thin sectioning, staining, critical point drying, sputter coating, microscope operation, photography, and dark room procedures. Students will complete a project of their choosing to demonstrate their ability to use their new skills. 2 LECT., 4 LAB HR./WK., PLUS 3 HR. TBA; 4 CR.

V8101: Mathematical Biology: 3 LECT. HR./WK.; 3 CR.

V8201: Biostatistics I: Univariate statistics of biological systems (theory and application). Topics include: probability, descriptive statistics, correlation, analysis of variance, and regression. Prereq: permission of instructor. 3 LECT., 6 LAB HR./WK.; 6 CR.

V9100: Colloquium: Recent developments and trends in the field of biology. Required of all candidates for the M.A. degree. 2 HR./WK.; 1 CR./SEM.

V9200: Tutorial 1-4 CR. V9201: Advanced Study Study in an area where formal course work is not given. Subject matter may vary from assigned current readings in a specialized area with reports to special laboratory or field work. Prereq: permission of instructor. 1-4 CR.

V9302: Molecular Biology Journal Club 1 CR. V9303: Seminar Special Topics Specialized seminars in diverse fields, depending upon the needs of specific students. 2 HR./WK., PLUS CONF.; 3 CR. GRADUATE

COURSES OPEN TO UNDERGRADUATES The following graduate courses are available to undergraduate students who have completed the appropriate prerequisites. Permission to take these courses must be obtained from the Biology Department Graduate Deputy Chairman and course instructor.

V0901/02: Population Genetics Lecture, Laboratory

V4103/04: Radiation Biology Lecture, Laboratory

V2800: Field Studies in Animal Behavior

V5303: Molecular Basis of Development

V6003/04: Community Ecology Lecture, Laboratory

V6005/06: Population Ecology

V6107/08: Marine Microbiology Lecture, Laboratory

V7200: Biological Electron Microscopy

V8101: Mathematical Biology


V2403: Animal Behavior I

V2407: Animal Behavior II

Contact Information

Tadmiri Venkatesh

Marshak Science Building
Room 526
160 Convent Avenue
New York, NY  10031

p:  212.650.6800
f:  212.650.8585